Epigenetics introduces additional gene regulation complexity beyond DNA sequence. This session will concentrate on DNA methylation, histone modification, and chromatin remodeling in plants and focus on their functions in development, stress response, and transgenerational inheritance. Epigenetic memory, as experts discuss it, refers to how plants "remember" environmental stresses and respond more efficiently in later generations. Case studies will illustrate how epigenetics controls flowering time, seed development, and resistance to pathogens. The latest developments in epigenomic mapping technologies and CRISPR-based epigenome editing will be discussed. The session will also discuss the application of epigenetics to plant improvement and sustainable agriculture. Through the revelation of how heritable gene expression changes influence plant behavior, the participants will understand this rapidly evolving field that links molecular biology, ecology, and evolution.