Epigenetic control indicates heritable changes in gene expression without modifying DNA sequences. DNA methylation, histone modifications, and chromatin remodeling will be discussed as central players in the regulation of gene activity in this session. Participants will be taught how non-coding RNAs impinge on epigenetic regulation. The session accentuates epigenetics in plant development, environmental adaptation, and transposon regulation. Researchers will provide case studies on stress memory, with plants "remembering" previous stresses for enhanced resilience. Participants will learn how epigenetics adds an extra control over and above genetics. The session will also address cutting-edge epigenomic methods for methylation and histone mapping. Applications include taking advantage of epigenetic regulation for plant breeding and stress tolerance. The session highlights the role of epigenetic plasticity in allowing quick adaptation to variable environments. Participants will also learn about the evolutionary relevance of epigenetics. The session bridges molecular biology, systems biology, and applied agriculture, giving a complete picture of epigenetic regulation in crops.